Paper 2, Section I, B

Mathematical Biology
Part II, 2011

A population with variable growth and harvesting is modelled by the equation

ut+1=max(rut21+ut2Eut,0)u_{t+1}=\max \left(\frac{r u_{t}^{2}}{1+u_{t}^{2}}-E u_{t}, 0\right)

where rr and EE are positive constants.

Given that r>1r>1, show that a non-zero steady state exists if 0<E<Em(r)0<E<E_{m}(r), where Em(r)E_{m}(r) is to be determined.

Show using a cobweb diagram that, if E<Em(r)E<E_{m}(r), a non-zero steady state may be attained only if the initial population u0u_{0} satisfies α<u0<β\alpha<u_{0}<\beta, where α\alpha should be determined explicitly and β\beta should be specified as a root of an algebraic equation.

With reference to the cobweb diagram, give an additional criterion that implies that α<u0<β\alpha<u_{0}<\beta is a sufficient condition, as well as a necessary condition, for convergence to a non-zero steady state.